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Synecology of the white-oak (Quercus garryana Douglas) woodlands of the Willamette Valley, Oregon

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Title Synecology of the white-oak (Quercus garryana Douglas) woodlands of the Willamette Valley, Oregon
Names Thilenius, John F., 1931- (creator)
Chilcote, William W. (advisor)
Date Issued 1963-10-14 (iso8601)
Note Graduation date: 1964
Abstract Quercus garryana dominated plant communities are found in
the interior coastal valleys and on foothills from southeastern
Vancouver Island, British Columbia, south to San Francisco,
California. They occur as savannas, intermixed with a variety of
conifers, and in almost pure stands. The diversity of habitats
occupied by the species and the variety of vegetation associated
with it provide an opportunity for a basic synecological investigation
of plant communities having a common, important member. The
study was designed to describe the floristic composition and structure
of the Quercus garryana communities, to determine their
ecological relation to the physical environment, and clarify successional
status. It provides a framework for future autecological
investigations of the component species and is directed at furnishing
a fuller understanding of the synecology of an important segment of
the vegetation of western Oregon.
Two levels of sampling were used during the study. The first
was a rapidly applied, qualitative method. This reconnaissance
technique was designed to provide an assessment of dominance, size-class distribution, and composition of the species comprising the
vegetation. Information on influential habitat factors was also obtained.
Sampling at this level enabled the examination of many
stands, provided a record of each stand, and served as the basis for
the selection of stands for more quantitative sampling.
One hundred forty stands were examined during the reconnaissance
phase of the study. Subjective evaluation of these stands supported
by an analysis using marginal punch cards determined that
only a few species were dominating the tall shrub and low shrub-herbaceous
layers of the understory and that certain combinations of
these dominant species were being repeated geographically through
the study area.
The second sampling phase was based on the reconnaissance
information and was designed to provide a quantitative record of the
species complexes or plant communities delineated during reconnaissance.
This method was applicable by a single investigator and
measured the dominance, frequency, size distribution, and density of
tree species; and the percent coverage and percent frequency of shrubs, herbs, and grasses. Soil profile descriptions were made in
each stand where the vegetation was quantitatively sampled. Salient
features of the physical environment were also recorded as were
indications of past and present land uses.
Forty seven stands were sampled. Quercus garryana was the
cverstory tree dominant in all of these stands. The understory
pecies were found to fall into four major communities. From
mesic to xeric these were: the Corylus cornuta/Polystichum
munitum community, the Prunus avium/Symphoricarpos albus community,
the Amelanchier alnifolia/Symphoricarpos albus community,
and the Rhus diversiloba community. These communities were
named for the species usually dominating the tall shrub and low
shrub layers. Seven soil series served as the substrate of these
communities; Steiwer, Carlton, Peavine, Dixonville, Nekia, Olympic,
and Amity. The understory plant communities of the Quercus
forest show considerable variation and intergradation. Changes are
a matter of shifts in species dominance rather than alterations in
species composition. The absence of environmental extremes in the
Willamette Valley and heavy present and historical land use have
increased this intergradation.
The presence of large, open-form Quercus garryana trees
surrounded by smaller forest-form trees indicates that the present
Quercus forests have developed from a savanna. The cause of this
change in gross physiognomy is probably the control of the repeated
ground fires which swept the pre-settlement savanna. Mature Quercus
trees are not harmed by ground fires, but such fires would tend
to keep dense reproduction from occurring.
The present abundance of Rhus diversiloba in the under story
may be directly related to heavy livestock grazing. The interconnected
root system of ground cover and liana-form Rhus provides
the species with a grazing resistance mechanism. The liana Rhus is
out of reach of grazing animals. Photosynthate transferred from the
liana to the ground cover plants would aid the latter in retaining vigor
even under heavy grazing pressure. Thus it would have an advantage
over other plants that were also being grazed, eventually
becoming the ground layer dominant.
Both Pseudotsuga menziesii and Acer macrophyllum appear to
be successional to Quercus garryana. Acer is better adapted to
mesic sites than Quercus. Pseudotsuga will succeed Quercus on
less mesic sites. The successional trends are promoted by livestock
grazing which opens up the ground layer and facilitates seedling
establishment especially for Pseudotsuga. On some sites Prunus
avium, an introduced species, is becoming part of the overstory canopy. It reproduces vigorously in its own shade and will become
an important member of the Quercus forest in the future. Quercus
garryana reproduction was more abundant on drier, exposed sites
and the species seems to be able to perpetuate itself on these
locations.
Genre Thesis/Dissertation
Topic Oregon oak -- Ecology -- Oregon -- Willamette River Valley
Identifier http://hdl.handle.net/1957/48442

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